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coli bacterial culture, which was first heat-inactivated (60 °C O/N), at 2000 × g for 10 min, and then concentrated two-fold in M9 plus cholesterol (5 μg mL −1 )

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Intestinal enterocytes absorb heme iron directly via haem carrier protein 1 (HCP1), while non-heme iron is transported by divalent metal transporter 1 (DMT1) after were reduced by reductase such as duodenal cytochrome b (Dcytb) to ferrous ions (Fe 2+ ) (Fig